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This course builds a coherent foundation in cell biology by connecting cell structure to function across molecular, organellar, and systems scales. Topics include cell theory; prokaryotic versus eukaryotic organization and evolutionary origins; macromolecules and enzyme principles; membrane structure, dynamics, and transport; the endomembrane system and organelle roles; bioenergetics in mitochondria (and chloroplasts overview); information flow from DNA to RNA to protein with protein targeting; mutations and multilayer gene regulation; signaling networks and feedback; the cell cycle, division, apoptosis, and cancer links; cytoskeleton-based transport and motility; cell interactions in tissues; and essential experimental methods and data interpretation.