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This course develops an integrative, research-oriented framework for explaining how and why cognitive abilities evolve across taxa, with special attention to humans. Core coverage includes evolutionary forces (selection, drift, mutation, constraint), adaptation vs exaptation, and Tinbergen’s four questions linking mechanism, development, function, and phylogeny. Methods emphasize comparative cognition task validity, phylogenetic comparative inference, and socio-ecological/behavioral-ecology models. Competing hypotheses (e.g., Social Brain, Machiavellian, Cultural Intelligence), gene–culture coevolution, and neuroscience/genetic evidence are evaluated alongside paleoanthropological and archaeological constraints. The course culminates in designing discriminative tests and synthesizing falsifiable evolutionary accounts.